As the script evolved over the centuries, the characters became more complex. Specifically, around 600 CE, flourishes and exaggerations of the basic strokes began to appear in more manuscripts. Ascenders and descenders were the first major alterations, followed by twists of the tool in the basic stroke and overlapping. By the time the more compact minuscule scripts awoke circa 800 CE, some of the evolved uncial styles formed the basis for these simplified, smaller scripts. Uncial was still used, particularly for copies of the Bible, tapering off until around the 10th century.
The word, uncial, is also sometimes used to refer to manuscripts that have been scribed in uncial, especially when differentiating from those which have been penned with minuscule. Some of the most noteworthy Greek uncials are:
Modern calligraphy usually teach a form of evolved Latin based uncial hand that would probably be best compared to the later, 7th to 10th century examples, though admittedly, the variations in Latin uncial are much wider and less rigid than Greek. Modern uncial has borrowed heavily from some of the conventions found in more cursive scripts, using flourishes, variable width strokes, and on occasion, even center axis tilt.