The Dart rises on Dartmoor, as two separate branches (the East Dart and West Dart), which join at Dartmeet. The paths along these rivers offer very attractive walking, and there are several small waterfalls. The rivers are crossed by a number of clapper bridges, notably at the hamlet of Postbridge.
After leaving the moor, the Dart flows southwards past Buckfast Abbey and through the towns of Buckfastleigh, Dartington and Totnes. From Totnes it becomes tidal, and there are no bridges below this point. The Dart estuary is a large ria and is popular for sailing. The towns of Kingswear and Dartmouth are on the east and west sides of the estuary, and are linked by a vehicle ferry.